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1. The Skull Spiders are part of the natural landscape of Okoto. This my be an "argument from ignorance" thing going. But somehow I doubt that the MoUP has the power to give life, only to corrupt life that exists (as witnessed with Makuta). So IMO the spideys would have to exist before the Ekimu incident. 2. If a creature (or perhaps even a villager) dies, it comes back as a Skull Spider. This sounds rather weird, but if a few small "Rahi" die and become Skull Spiders every so now and again, they wouldn't amass a huge army of them, presuming that a few of them get destroyed/eaten every so often. But if there was a massive shockwave to kill of a huge number of them at once, then a huge army of them could appear and become a problem. 3. Larger creatures, upon their death, become the Lord of Skull Spiders and similar villain-type-things. Given the normal rate of death, there probably would only be one of them at a time to direct the Spiders and keep them away from the villages. Now there are a bunch of them, and they could be mad at the villagers for causing their death via mask. And they want the masks for themselves to avoid something like that from happening again. I'm sure you can find a hole or two in this spiderweb. Go!
The Verdelisks are a species of Protector-sized dragon-like reptiles that are native to Okoto's jungle region (though a few sometimes wander into the swamps of the Region of Water). Their wings are too small for them to fly; at best, they can glide for short distances, which is enough to allow them to hop from tree to tree. When in large open spaces, the Verdelisks use their wings for short hops, giving them a notable speed boost. The villagers of the jungle have tamed a few Verdelisks and sometimes use them to carry goods as part of a caravan. These individuals wear special harnesses with containers on them. What they gain in storage, they lose in speed, as they have difficulty using their wings while wearing the harnesses. Other pictures: Top picture-larger All fours Gliding back With harness Opened container Upright with harness